Isoetes engelmannii is a species of aquatic plant in the family Isoetaceae. It is referred to by the common names Engelmann's quillwort or Appalachian quillwort (not to be confused with the newly described Isoetes appalachiana), and is the most widely distributed species of its genus in eastern North America. Its range extends from Ontario in the north, south to Florida and west Arkansas and Missouri. It can be found from April to October in temporary pools, bogs, marshes, stream edges, swamps and along wet roadsides.
No narrative description available for this taxon yet.
A DNA barcode is a short, standardised stretch of genes that works like a fingerprint — enough to tell one species from another. Below is the molecular trace Isoetes engelmannii has left across the world's sequence archives.
At a glance
DNA specimens16
Marker genes6
GenBank sequences10
eDNA detections33
Countries2
The DNA barcodea real sequence read deposited for this species
★ the standard DNA barcode for this group — the short region actually read to tell this species apart. The rest are extra genes sequenced along the way.
★COI-5P★rbcL5★rbcLa★ITS5★ITS2NBC-COI-5P
animal barcodeplant barcodefungal barcodemarker
06Genome at a glanceCCDB · GoaT · NCBI
The complete instruction manualIsoetes engelmannii carries — its genome. We read it from three angles — how big it is, how the DNA is packed into chromosomes, and how completely it has been sequenced — and explain how to read each value as you go.
Genome sizehow big the whole instruction manual is
Genome size≈640 967 017 bp assembly estimate
Measured in base pairs (bp) — the individual letters of DNA (human ≈ 3.2 Gb, a bacterium a few million). The chart places this genome on a logarithmic scale — each step to the right is ten times bigger — among reference organisms. Across species a bigger genome loosely tracks with larger cells, slower growth and lower-energy lifestyles (powered flight favours small genomes) — yet it does not imply more genes or a more advanced organism (the long-standing C-value paradox).
BACTERIUM Carsonella ruddii0.00016 Gb
FUNGUS0.04 Gb
INSECT0.25 Gb
THIS GENOME Isoetes engelmannii0.64 Gb
HUMAN3.2 Gb
WHEAT17 Gb
FERN Tmesipteris160.45 Gb
Chromosomes & ploidyhow the DNA is packaged
2n is the full chromosome count in a normal body cell; n is a gamete (egg or sperm), which carries half. Ploidy is how many complete chromosome sets each cell holds — 2× (diploid) is typical for animals, while higher levels (polyploidy) are common in plants. Click any value below to see the underlying records and sources.
CCDB · ipcn-api-dl — Winstead, J. E. & L. J. Musselman. 1994. Ecophenic patterns of Isoetes engelmannii in a roadside and pond habitat of south central Kentucky [abstract]. ASB Bull. 41: 123.
CCDB · ipcn-api-dl — Musselman, L. J., R. D. Bray & D. A. Knepper. 1997. Isoetes @xcarltaylori (Isoetes acadiensis @x Isoetes engelmannii), a new interspecific quillwort hybrid from the Chesapeake Bay. Canad. J. Bot. 75: 301–309.
CCDB · ipcn-api-dl — Musselman, L. J., D. A. Knepper, R. D. Bray, C. A. Caplen & C. Ballou. 1995. A new Isoetes hybrid from Virginia. Castanea 60(3): 245–254.
CCDB · ipcn-api-dl — Barr, A. D. & L. J. Musselman. 1997. New hybrids in the genus Isoetes (quillworts) in the southeastern United States. (Abstract). ASB Bull. 44(2): 119–120.
CCDB · ipcn-api-dl — Brunton, D. F. & D. M. Britton. 1996. Taxonomy and distribution of Isoetes valida. Amer. Fern J. 86(1): 16–25.
CCDB · ipcn-api-dl — Montgomery, J. D. & W. C. Taylor. 1994. Confirmation of a hybrid Isoetes from New Jersey. Amer. Fern J. 84: 115–120.
n 111×CCDB · book-ipcn66
CCDB · book-ipcn66 — Wagner & Wagner 1966
Sequencing statusassembly quality — how far to trust these numbers
Assembly level tells you how finished the sequence is — from fragmented contigs, through scaffolds, up to a full chromosome-level assembly. BUSCO % estimates completeness: the share of genes expected to be present that were actually found. These describe the data quality, not the organism.
Assembly levelScaffold
Completeness69.4% BUSCO
07Deep time~3.33 Ma lineage
How far back this lineage goes — and how we know. Everything here is measured in Ma, short for “mega-annum”: millions of years ago. The chart reads left to right like a calendar of the Earth, from the deep past on the left to today at the right edgetop to bottom like a core drilled through the Earth, from the deep past at the top down to today at the bottom.
At a glance
DNA clock origin3.33 Ma TimeTree
When this lineage existed
How to read this: the coloured strip along the bottomdown the left is the geological calendar — the standard epochs (Pliocene, Pleistocene…) every museum uses, shown so you can see which chapter of Earth's history this lineage lived in. This lineage is a young one, so the strip is zoomed in to epochs — the finer subdivisions inside a period. The orange marker is the DNA clock: DNA accumulates mutations at a roughly steady rate, so comparing this species' DNA with its relatives estimates when the lineage split off — independently of any fossil.
DNA clock origin
08Occurrence & distribution
Record type709 records
Wild obs. + sensor11
Museum / vouchered698
Range
Area of Occupancy AOO1 972 km²
Wildobservation + sensor
Human sightings and records, or camera-trap / sensor detections — someone (or a device) saw or captured the species in the wild.
Coordinate accuracy50% within 1 km
≤100 m 5>10 km 5
10 georeferenced · 1 without coordinates
Open the mapobservation + sensor11
Museum / Voucheredphysical evidence
Backed by a physical specimen — a herbarium sheet, sample or voucher held in a collection. “Vouchered” means supported by material evidence, not just an observation.
Coordinate accuracy35% within 1 km
≤100 m 17≤1 km 151≤10 km 286>10 km 25
479 georeferenced · 219 without coordinates
Open the institutions mapphysical evidence698
Wildobservation + sensor
Human sightings and records, or camera-trap / sensor detections — someone (or a device) saw or captured the species in the wild.
10Collections & institutions
Holding institutions40 of 61 geolocated
Institutions and collections holding physical, vouchered specimens of this species — click a row to fly to it on the map.
Institution
Specimens
Philadelphia, US
133
Chapel Hill, US
56
Chongqing Museumlocation not on record
46
Minia, EG
37
Bronx, US
29
Norfolk, US
29
Knoxville, US
27
Philadelphia, US
23
University of New Hampshirelocation not on record
20
Florida Museum of Natural History- Zoology, Paleontology & Paleobotanylocation not on record
17
Burlington, US
16
Durham, US
11
Bangkok, TH
11
McWane Science Centerlocation not on record
11
University of Stellenboschlocation not on record
11
GAlocation not on record
10
Chicago, US
9
New Brunswick, US
8
Williamsburg, US
8
Ann Arbor, US
8
LDlocation not on record
7
Staten Island, US
6
Appalachian State Universitylocation not on record
5
Bloomington, US
5
Allentown, US
5
BAYLUlocation not on record
4
Saint Louis, US
4
Columbia, US
4
Auckland, NZ
4
WTUlocation not on record
3
Tampa, US
3
San Angelo, US
3
Morgantown, US
3
Fort Worth, US
3
Lord Fairfax Community Collegelocation not on record
3
Logan, US
2
New Haven, US
2
Maryland Department of Natural Resourceslocation not on record
2
Tuscaloosa, US
2
Denver, US
2
St. Paul, US
2
Millersville, US
2
Valdosta State Universitylocation not on record
2
Oswego, US
2
Clemson, US
2
Tall Timbers Research Stationlocation not on record
1
James F. Matthews Center for Biodiversity Studieslocation not on record
1
Macomb, US
1
Istituto Agrario Castelnuovolocation not on record
1
Fairfax, US
1
Pocatello, US
1
University of Tennessee at Chattanoogalocation not on record
1
Québec, CA
1
Western Carolina Universitylocation not on record
1
University of North Carolina at Pembrokelocation not on record
1
DOI/NPS, Mississippi National River & Recreation Arealocation not on record
1
Meguro Parasitological Museumlocation not on record
1
Mississippi State, US
1
Austin, US
1
Flagstaff, US
1
Madison, US
1
61 institutions · 618 of 698 vouchered records shown · 80 without an institution code
09Environmental DNA33 detections
Where the DNA of Isoetes engelmannii was picked up in samples of water, soil or air — nobody saw the organism, only its DNA left behind. A trace is a clue that the species was near, not a confirmed sighting.
Signal
Detections DNA found33
Studies independent surveys2
Countries1
Signal confidence: moderateweighed across independent studies, places & mapped detections
Where its DNA was found
0 of 33 detections have coordinates
Open the map1 country0
How strong is each trace?
DNA read depthRead counts were not reported for this species — the map shows presence only, not how strong each trace was.
Modelled climatemodelled
−15°Ctemperature across detection sites+40°C
Temperature median9.60 °C 8.30–11.6
Seasonal swing summer↔winter16.8 °C
Max temp (day)12.1 °C 10.6–14.2
Min temp (night)6.80 °C 4.10–7.30
Precipitation143 mm/mo 94.3–177
Air humidity63.4 % 60.9–66.9
Moisture balance42.2 mm/mo -6.40–125
Vapour deficit440 Pa 381–539
Wind speed3.60 m/s 3.10–5.00
Cloud cover50.2 % 49.0–52.8
CHELSA 1981–2010, ~9 km grid, at location & month of 27 detection points · median with p10–p90 · reflects where sampling happened, not only the true niche
How to read this: each dot is one detection of this species' DNA in an environmental sample. The confidence meter weighs how many independent studies and places back up the signal — one detection in one study is a hint; many across several studies is solid. Records dated before 2008 (when eDNA methods began) are treated as likely mislabeled and left off the map.
WikipediaWikimedia Foundation3 languages↗
BioWikiNetmultilingual Wikipediamultilingual↗
GBIFGlobal Biodiversity Information Facility709 records↗
ENAEuropean Nucleotide Archive · EMBL-EBI33 eDNA detections↗
BOLD SystemsCentre for Biodiversity Genomics16 specimens↗
NCBIUS National Library of Medicinesequences↗
Open Tree of LifeOpenTreephylogeny backbone↗
CCDBChromosome Counts DB · Tel Aviv U.genome & karyotype↗
GoaTGenomes on a Tree · Sangergenome & karyotype↗
WikidataWikimedia Foundationstructured facts↗
GLoBIGlobal Biotic Interactionsbiotic interactions↗