Dryopteris carthusiana
(Vill.) H.P.Fuchs · speciesAt a glance
Sources12 archives
Databases and archives Dryopteris carthusiana's data was compiled from.
WikipediaWikimedia Foundation10 languages↗
BioWikiNetmultilingual Wikipediamultilingual↗
GBIFGlobal Biodiversity Information Facility398 534 records↗
ENAEuropean Nucleotide Archive · EMBL-EBI32 eDNA detections↗
BOLD SystemsCentre for Biodiversity Genomics44 specimens↗
NCBIUS National Library of Medicinesequences↗
LOTUSNatural Products (Wikidata)compounds↗
GRIN TaxonomyUSDA-ARSdistribution & uses↗
Open Tree of LifeOpenTreephylogeny backbone↗
CCDBChromosome Counts DB · Tel Aviv U.genome & karyotype↗
GoaTGenomes on a Tree · Sangergenome & karyotype↗
GLoBIGlobal Biotic Interactionsbiotic interactions↗Every layer below draws on the sources above — open one to explore it, or use ← → to move between tabs.
Dryopteris carthusiana is a species of fern native to damp forests throughout the Holarctic Kingdom. It is known as the narrow buckler-fern in the United Kingdom, and as the spinulose woodfern in North America.USDA PLANTS Profile It is a tetraploid of hybrid origin, one parent being Dryopteris intermedia, known in North America as the intermediate wood fern, and an unknown, apparently extinct species dubbed Dryopteris semicristata, which is also the presumed parent of the hybrid-origin Dryopteris cristata. Showing sori and leaflet shape This fern is often confused with several other wood fern species, including D. intermedia, D. campyloptera, and D. expansa. It especially extensively shares the range of D. intermedia, but the two may be distinguished by the innermost pinnule on the bottom side of the bottom pinna: this pinnule is longer than the adjacent pinnules in D. carthusiana, but shorter or even in D. intermedia. D. carthusiana is a sub-evergreen species, its fronds surviving mild winters but dying back in harsh winters. A nature print of Dryopteris carthusiana
No narrative description available for this taxon yet.
Size & morphology6
Life cycle & reproduction10
Diet & foraging2
Habitat & environment9
Physiology & chemistry6
Other traits1
Compounds documented for Dryopteris carthusiana across natural-product and food-composition databases — not just the ~150 nutrients on a classic label ("nutritional dark matter").
Compound class profile5 classes
Documented compounds22 total
| Compound | Class | Amount | Source |
|---|---|---|---|
| (3R,3'R,13-cis)-beta,beta-Carotene-3,3'-diol | present | LOTUS | |
| 1-Heptadecene | present | LOTUS | |
| alpha-Carotene | present | LOTUS | |
| Behnic acid | present | LOTUS | |
| beta-Carotene | present | LOTUS | |
| Cryptoxanthin | present | LOTUS | |
| Hexadecane | present | LOTUS | |
| Hexadecenoic acid | present | LOTUS | |
| Linonelic acid | present | LOTUS | |
| Lutein | present | LOTUS |
A DNA barcode is a short, standardised stretch of genes that works like a fingerprint — enough to tell one species from another. Below is the molecular trace Dryopteris carthusiana has left across the world's sequence archives.
At a glance
★ the standard DNA barcode for this group — the short region actually read to tell this species apart. The rest are extra genes sequenced along the way.
The complete instruction manual Dryopteris carthusiana carries — its genome. We read it from three angles — how big it is, how the DNA is packed into chromosomes, and how completely it has been sequenced — and explain how to read each value as you go.
Genome sizehow big the whole instruction manual is
Measured in base pairs (bp) — the individual letters of DNA (human ≈ 3.2 Gb, a bacterium a few million). The chart places this genome on a logarithmic scale — each step to the right is ten times bigger — among reference organisms. Across species a bigger genome loosely tracks with larger cells, slower growth and lower-energy lifestyles (powered flight favours small genomes) — yet it does not imply more genes or a more advanced organism (the long-standing C-value paradox).
Chromosomes & ploidyhow the DNA is packaged
2n is the full chromosome count in a normal body cell; n is a gamete (egg or sperm), which carries half. Ploidy is how many complete chromosome sets each cell holds — 2× (diploid) is typical for animals, while higher levels (polyploidy) are common in plants. Click any value below to see the underlying records and sources.
2n 16433×GoaT · DTOL Flowering Plants Estimates Kew · GoaT · Kew Plant DNA C-values Database · CCDB · brit-fl +6
2n 1841×CCDB · Cave1959
n 8211×CCDB · ipcn-api-dl · CCDB · book-ipcn66 · CCDB · book-ipcn67-71 +5
n 411×CCDB · Cave1963
tetraploid1×GoaT · Kew Plant DNA C-values Database
How far back this lineage goes — and how we know. Everything here is measured in Ma, short for “mega-annum”: millions of years ago. The chart reads left to right like a calendar of the Earth, from the deep past on the left to today at the right edgetop to bottom like a core drilled through the Earth, from the deep past at the top down to today at the bottom.
At a glance
When this lineage existed
How to read this: the coloured strip along the bottomdown the left is the geological calendar — the standard epochs (Pliocene, Pleistocene…) every museum uses, shown so you can see which chapter of Earth's history this lineage lived in. This lineage is a young one, so the strip is zoomed in to epochs — the finer subdivisions inside a period. The orange marker is the DNA clock: DNA accumulates mutations at a roughly steady rate, so comparing this species' DNA with its relatives estimates when the lineage split off — independently of any fossil.
Record type398 534 records
Origin
Range
Wildobservation + sensor
Human sightings and records, or camera-trap / sensor detections — someone (or a device) saw or captured the species in the wild.
Museum / Voucheredphysical evidence
Backed by a physical specimen — a herbarium sheet, sample or voucher held in a collection. “Vouchered” means supported by material evidence, not just an observation.
Cultivated / Captivenot free-living
A living individual in a botanical garden, zoo or nursery — cultivated or kept, not free-living.
Wildobservation + sensor
Human sightings and records, or camera-trap / sensor detections — someone (or a device) saw or captured the species in the wild.
Holding institutions136 of 268 geolocated
Institutions and collections holding physical, vouchered specimens of this species — click a row to fly to it on the map.
| Institution | Specimens |
|---|---|
| Uniwersytet Wrocławskilocation not on record | 29 266 |
| Helsinki, FI | 905 |
| Moscow State Universitylocation not on record | 616 |
| Olocation not on record | 602 |
| LDlocation not on record | 581 |
| BFLlocation not on record | 502 |
| St. Paul, US | 494 |
| Chongqing Museumlocation not on record | 474 |
| Madison, US | 430 |
| Institute of Plant and Animal Ecology Ural Branch of the Russian Academy of Scienceslocation not on record | 429 |
| Provincia di Livornolocation not on record | 421 |
| MeiseBGlocation not on record | 290 |
| Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)location not on record | 273 |
| Oulu, FI | 271 |
| Ann Arbor, US | 225 |
| KMNlocation not on record | 209 |
| SLU Artdatabankenlocation not on record | 198 |
| Museo civico La Terra e l'Uomo di Crocetta del Montellolocation not on record | 193 |
| Uniwersytet Rolniczy im. Hugona Kołłątaja w Krakowielocation not on record | 191 |
| Montréal, CA | 184 |
| University of Silesia in Katowicelocation not on record | 181 |
| Bergen, NO | 175 |
| GBS RAN - Glavny Botanichesky Sad Rossijskoj Akademii Nauklocation not on record | 168 |
| Zürich, CH | 166 |
| Zürich, CH | 152 |
| DBF-NHMDlocation not on record | 149 |
| Trondheim, NO | 148 |
| Salzburg, AT | 141 |
| Turku, FI | 133 |
| Uniwersytet Śląski w Katowicachlocation not on record | 132 |
| IFR-DNFlocation not on record | 131 |
| Oskarshamn, SE | 125 |
| Burlington, US | 119 |
| McWane Science Centerlocation not on record | 112 |
| South Kensington, GB | 106 |
| Acadia Universitylocation not on record | 102 |
| MAlocation not on record | 100 |
| Bloomington, US | 98 |
| Green Bay, US | 94 |
| Görlitz, DE | 90 |
| University of Alberta Museumslocation not on record | 86 |
| Minia, EG | 81 |
| Jyväskylä, FI | 78 |
| Toronto, CA | 78 |
| Tartu, EE | 77 |
| Durham, US | 77 |
| BIO-UNIPIlocation not on record | 75 |
| Bronx, US | 69 |
| WINlocation not on record | 66 |
| Tallinn, EE | 65 |
| Université Lavallocation not on record | 64 |
| Udmurt State Universitylocation not on record | 62 |
| Chapel Hill, US | 61 |
| Berlin, DE | 59 |
| Kuopio, FI | 58 |
| Vancouver, CA | 57 |
| BRNUlocation not on record | 45 |
| Tyumen State Universitylocation not on record | 44 |
| SZUlocation not on record | 43 |
| Dresden, DE | 43 |
| GJOlocation not on record | 42 |
| Irkutsk State Universitylocation not on record | 42 |
| TROMlocation not on record | 40 |
| IPDNlocation not on record | 40 |
| Wlocation not on record | 40 |
| Chicago, US | 39 |
| Philadelphia, US | 38 |
| WTUlocation not on record | 38 |
| Montréal, CA | 37 |
| University of Calgarylocation not on record | 36 |
| University of Stellenboschlocation not on record | 36 |
| Allentown, US | 35 |
| Saint John, CA | 34 |
| Philadelphia, US | 33 |
| Québec, CA | 32 |
| Uniwersytet Warszawskilocation not on record | 32 |
| Bern, CH | 32 |
| València, ES | 32 |
| Vitoria, ES | 32 |
| Missoula, US | 32 |
| Museum Ludovicae Ulricae, Zoology Institute of the University of Uppsalalocation not on record | 31 |
| Tampere, FI | 29 |
| Victoria, CA | 28 |
| Pullman, US | 28 |
| Moscow, US | 28 |
| Frankfurt am Main | 27 |
| Institute of Biology Ufa Scientific Centre of Russian Academy of Scienceslocation not on record | 27 |
| Tula State Lev Tolstoy Pedagogical Universitylocation not on record | 25 |
| San Sebastián, ES | 24 |
| Uppsala, SE | 24 |
| Universidad del Pais Vasco (UPV/EHU)location not on record | 24 |
| Salamanca, ES | 24 |
| Morgantown, US | 21 |
| Entomological Society of Latvialocation not on record | 21 |
| Saint Louis, US | 21 |
| CJBGlocation not on record | 20 |
| Tula State Universitylocation not on record | 20 |
| Wellington, NZ | 19 |
| University of New Hampshirelocation not on record | 19 |
| TUR-Alocation not on record | 19 |
| Ernst-Moritz-Arndt-Universitat Greifswaldlocation not on record | 19 |
| Frauenfeld, CH | 19 |
| Barcelona, ES | 19 |
| GZUlocation not on record | 19 |
| Millersville, US | 19 |
| Tampa, US | 19 |
| Philadelphia, US | 19 |
| Mlocation not on record | 18 |
| BDBClocation not on record | 17 |
| Santiago de Compostela, ES | 17 |
| Auckland, NZ | 17 |
| Podgorica, ME | 16 |
| Kandalaksha State Nature Reservelocation not on record | 16 |
| Natural History Museum Rotterdamlocation not on record | 15 |
| Fort Worth, US | 15 |
| Chelyabinsk State Universitylocation not on record | 15 |
| Taipei, TW | 14 |
| Madrid, ES | 14 |
| New Haven, US | 13 |
| Corvallis, US | 13 |
| PRClocation not on record | 12 |
| John May Museum of Natural Historylocation not on record | 12 |
| Lincoln, US | 12 |
| Porvoo, FI | 12 |
| Karlsruhe, DE | 12 |
| NMBU:MINAlocation not on record | 12 |
| Norfolk, US | 11 |
| Kurgan State Universitylocation not on record | 11 |
| PrivatHerblocation not on record | 11 |
| Rovaniemi, FI | 10 |
| Jaén, ES | 10 |
| Stockholm, SE | 10 |
| University of Wisconsinlocation not on record | 9 |
| Austin, US | 9 |
| Whitewater, US | 9 |
| Tilburg, NL | 9 |
| University of Copenhagen, Natural History Museum of Denmark, Herbarium Clocation not on record | 9 |
| US | 9 |
| College Park, US | 8 |
| Instytut Ochrony Przyrody Polskiej Akademii Nauklocation not on record | 8 |
| Lausanne, CH | 8 |
| Paris, FR | 8 |
| University of Winnipeglocation not on record | 8 |
| The State Museum-Preserve "The Kulikovo pole"location not on record | 7 |
| Royal Botanical Gardenslocation not on record | 7 |
| Science Museum of Minnesotalocation not on record | 7 |
| Tacoma, US | 7 |
| Williamsburg, US | 7 |
| Porrentruy, CH | 7 |
| Granada, ES | 7 |
| Appalachian State Universitylocation not on record | 7 |
| BSBIlocation not on record | 6 |
| Whitehorse, CA | 6 |
| EL PASO, US | 6 |
| Columbia, US | 6 |
| FLADlocation not on record | 6 |
| Fulda, DE | 6 |
| Boise, US | 6 |
| Chicago, US | 6 |
| Wuzhou, CN | 6 |
| New Brunswick, US | 6 |
| Urumqi, CN | 6 |
| Beijing, CN | 6 |
| Glarus, CH | 6 |
| UFPRlocation not on record | 5 |
| Oswego, US | 5 |
| Fairfax, US | 5 |
| Texas A&M Universitylocation not on record | 5 |
| Mount Pleasant, US | 5 |
| Prioksko-Terrasnyi Biosphere Reservelocation not on record | 5 |
| Forssa, FI | 5 |
| BClocation not on record | 4 |
| Fargo, US | 4 |
| Smithsonian Institutionlocation not on record | 4 |
| Institut und Museum fuer Geologie und Palaeontologielocation not on record | 4 |
| Provo, US | 4 |
| Chicago, US | 4 |
| Lord Fairfax Community Collegelocation not on record | 4 |
| Nagano City, JP | 3 |
| San Jose State University, Museum of Birds and Mammalslocation not on record | 3 |
| Yukon Universitylocation not on record | 3 |
| Siberian Federal Universitylocation not on record | 3 |
| Naturalis Biodiversity Centerlocation not on record | 3 |
| University of Guelph, OAC Herbariumlocation not on record | 3 |
| Chadron, US | 3 |
| Conservatoire botanique national du Bassin parisienlocation not on record | 3 |
| Adam Mickiewicz University in Poznańlocation not on record | 3 |
| The Tula Region State Institution of Culture "Tula Museum Association"location not on record | 3 |
| EEZA-CSIClocation not on record | 3 |
| Black Rock Forest Consortiumlocation not on record | 3 |
| Tempe, US | 3 |
| Mohonk Preservelocation not on record | 3 |
| UoB-IBlocation not on record | 3 |
| Riverside, US | 3 |
| Ypsilanti, US | 3 |
| Maryland Department of Natural Resourceslocation not on record | 3 |
| ASUlocation not on record | 2 |
| Namur, BE | 2 |
| Göteborg, SE | 2 |
| Boumlocation not on record | 2 |
| ZRC-SAZUlocation not on record | 2 |
| nbflocation not on record | 2 |
| Hobart, AU | 2 |
| Dekalb, US | 2 |
| Museo Entomologico de Leonlocation not on record | 2 |
| DULlocation not on record | 2 |
| Royal Alberta Museumlocation not on record | 2 |
| Museum of the Rockieslocation not on record | 2 |
| US | 2 |
| Museo Achille Folettolocation not on record | 2 |
| Florida Museum of Natural History- Zoology, Paleontology & Paleobotanylocation not on record | 2 |
| DOI/NPS, Little Rock Central High School National Historic Sitelocation not on record | 2 |
| Mississippi State, US | 2 |
| Bangkok, TH | 2 |
| Muzaffarabad, PK | 2 |
| Ivano-Frankove, UA | 2 |
| GAlocation not on record | 2 |
| H-Wolflocation not on record | 2 |
| CBDClocation not on record | 2 |
| Asheville, US | 2 |
| H-Adlerlocation not on record | 2 |
| Tsitsin Main Botanical Garden Russian Academy of Scienceslocation not on record | 2 |
| Decorah, US | 2 |
| Meguro Parasitological Museumlocation not on record | 2 |
| Samara National Research Universitylocation not on record | 2 |
| Gijón, ES | 2 |
| Tromso University Museumlocation not on record | 2 |
| H-AkrKGlocation not on record | 1 |
| Long Beach, US | 1 |
| H-Hohlalocation not on record | 1 |
| H-Dueringlocation not on record | 1 |
| MILlocation not on record | 1 |
| Christchurch, NZ | 1 |
| UGentlocation not on record | 1 |
| Pamplona, ES | 1 |
| NoColllocation not on record | 1 |
| Arequipa, PE | 1 |
| McGill University, Department of Biologylocation not on record | 1 |
| Keene State Universitylocation not on record | 1 |
| Macomb, US | 1 |
| Royal Botanic Gardens, Kewlocation not on record | 1 |
| Sion, CH | 1 |
| LfUlocation not on record | 1 |
| Tobolsk Complex Scientific Station of the Ural Branch of the Russian Academy of Scienceslocation not on record | 1 |
| Springfield, US | 1 |
| DOI/NPS, Greenbelt Parklocation not on record | 1 |
| McGill University, Herbariumlocation not on record | 1 |
| Research Collection of B. A. Bennettlocation not on record | 1 |
| Brookings, US | 1 |
| Museo civico di Storia naturale Giacomo Doria di Genova | Giacomo Doria Natural History Museum in Genoalocation not on record | 1 |
| San Angelo, US | 1 |
| Yugra State University Biological Collectionlocation not on record | 1 |
| Little Rock, US | 1 |
| H-Schuhwlocation not on record | 1 |
| Fribourg, CH | 1 |
| CASlocation not on record | 1 |
| JBRJlocation not on record | 1 |
| CSBSJUlocation not on record | 1 |
| Elikins, US | 1 |
| Canberra, AU | 1 |
| Alicante, ES | 1 |
| Brisbane, AU | 1 |
| Uniwersytet w Białymstokulocation not on record | 1 |
| Xiamen, CN | 1 |
| Bourges, FR | 1 |
| USP-IBlocation not on record | 1 |
| Canadian Department of Agriculturelocation not on record | 1 |
| München, DE | 1 |
Cultivated / Captivenot free-living
A living individual in a botanical garden, zoo or nursery — cultivated or kept, not free-living.
Where the DNA of Dryopteris carthusiana was picked up in samples of water, soil or air — nobody saw the organism, only its DNA left behind. A trace is a clue that the species was near, not a confirmed sighting.
Signal
Where its DNA was found
How strong is each trace?
Modelled climatemodelled
How to read this: each dot is one detection of this species' DNA in an environmental sample. The confidence meter weighs how many independent studies and places back up the signal — one detection in one study is a hint; many across several studies is solid. Records dated before 2008 (when eDNA methods began) are treated as likely mislabeled and left off the map.