Every layer below draws on the sources above — open one to explore it, or use ← → to move between tabs.
Traits & measurements1 literature sources4 traits
Occurrence & distributionGBIF · OBIS · PBDB587 records
Deep timePBDB · TimeTree39 fossils
02Traits & measurements
SourcesSeaLifeBase
Habitat & environment2
Habitat saltwateryes
Habitat zonebenthic
Other traits2
Depth max50 m
Depth min10 m
07Deep time23–2.58 Ma
How far back this lineage goes — and how we know. Everything here is measured in Ma, short for “mega-annum”: millions of years ago. The chart reads left to right like a calendar of the Earth, from the deep past on the left to today at the right edgetop to bottom like a core drilled through the Earth, from the deep past at the top down to today at the bottom.
At a glance
Fossil range23–2.58 Ma Oligocene, Miocene, Pliocene
Dated fossil finds39
Most finds around6.5 Ma 13 finds · Miocene
DNA clock origin32.3 Ma TimeTree
Ghost lineage9.27 Myr older than any fossil
StatusExtinct no longer alive anywhere
When this lineage existed
How to read this: the coloured strip along the bottomdown the left is the geological calendar — the standard epochs (Pliocene, Pleistocene…) every museum uses, shown so you can see which chapter of Earth's history this lineage lived in. This lineage is a young one, so the strip is zoomed in to epochs — the finer subdivisions inside a period. The solid bar is the fossil range: the span between the oldest and the youngest fossil that palaeontologists have assigned to Amusium papyraceum. Above itBeside it, the bars count how many dated finds fall in each slice of time; the tallest bar is labelled, and heights use a square-root scale so that thin slices stay visible next to rich ones. Read this as how well each stretch of time is preserved and studied — thick bars mean plenty of the right kind of rock and plenty of collectors, which is related to, but not the same as, how common it actually was. The orange marker is the DNA clock: DNA accumulates mutations at a roughly steady rate, so comparing this species' DNA with its relatives estimates when the lineage split off — independently of any fossil. Where the DNA reaches further back than the oldest fossil, the gap is hatched: the ghost lineage. It means the lineage was already out there, but has left us nothing we have dug up yet.
fossil range (PBDB) †dated finds per sliceDNA clock originghost lineage
How it livedPBDB
Environmentmarine
Life habitepifaunal
Dietsuspension feeder
Motilityactively mobile
Visionlimited
Compositionlow Mg calcite, aragonite
08Occurrence & distribution
Record type587 records
Wild obs. + sensor213
Museum / vouchered330
Fossil44
Origin
Native11
Range
Area of Occupancy AOO888 km²
Depth
0–200 m sunlit107
200–1000 m twilight2
1–4 km midnight0
>4 km abyssal0
median 55 m · max 274 m · 109 records with depth
Wildobservation + sensor
Human sightings and records, or camera-trap / sensor detections — someone (or a device) saw or captured the species in the wild.
Coordinate accuracy
no georeferenced coordinates · 213 records without
Open the mapobservation + sensor213
Museum / Voucheredphysical evidence
Backed by a physical specimen — a herbarium sheet, sample or voucher held in a collection. “Vouchered” means supported by material evidence, not just an observation.
Coordinate accuracy54% within 1 km
≤100 m 17≤1 km 4≤10 km 10>10 km 8
39 georeferenced · 291 without coordinates
Open the institutions mapphysical evidence330
Wildobservation + sensor
Human sightings and records, or camera-trap / sensor detections — someone (or a device) saw or captured the species in the wild.
10Collections & institutions
Holding institutions4 of 15 geolocated
Institutions and collections holding physical, vouchered specimens of this species — click a row to fly to it on the map.
Institution
Specimens
Washington, US
87
Florida Museum of Natural History- Zoology, Paleontology & Paleobotanylocation not on record
16
Cambridge, US
12
Texas Cooperative Wildlife Collectionlocation not on record
11
Florida Atlantic University, Harbor Branch Oceanographic Museumlocation not on record
2
New Haven, US
2
UM-RSMASlocation not on record
2
Natural History Museum of Utahlocation not on record
1
Alabama Museum of Natural Historylocation not on record
1
Sam Noble Oklahoma Museum of Natural Historylocation not on record
1
Instituto de Ciencias del Mar y Limnología, Unidad Académica Mazatlán, Universidad Nacional Autónoma de Méxicolocation not on record
1
Provincia di Livornolocation not on record
1
Chicago, US
1
North Carolina Museum of Natural Scienceslocation not on record
1
Paleontological Research Institutionlocation not on record
1
15 institutions · 140 of 330 vouchered records shown · 23 without an institution code